Occurrence Dot Map:
Data from 1409 occurrences (711 museum vouchers).
- 260 museum vouchers > 30 yrs.
- 451 museum vouchers < 30 yrs.
- 698 observations.
- 19 literature observations.
340 unique localities.

Collection/Observation Date Quartile Plot:

Collection/Observation by Hour:
10 record(s) from 12:00:00 AM to 1:00:00 AM 6 record(s) from 1:00:00 AM to 2:00:00 AM 0 records from 2:00:00 AM to 3:00:00 AM 0 records from 3:00:00 AM to 4:00:00 AM 0 records from 4:00:00 AM to 5:00:00 AM 0 records from 5:00:00 AM to 6:00:00 AM 0 records from 6:00:00 AM to 7:00:00 AM 0 records from 7:00:00 AM to 8:00:00 AM 1 record(s) from 8:00:00 AM to 9:00:00 AM 0 records from 9:00:00 AM to 10:00:00 AM 2 record(s) from 10:00:00 AM to 11:00:00 AM 0 records from 11:00:00 AM to 12:00:00 PM 0 records from 12:00:00 PM to 1:00:00 PM 0 records from 1:00:00 PM to 2:00:00 PM 1 record(s) from 2:00:00 PM to 3:00:00 PM 0 records from 3:00:00 PM to 4:00:00 PM 2 record(s) from 4:00:00 PM to 5:00:00 PM 0 records from 5:00:00 PM to 6:00:00 PM 0 records from 6:00:00 PM to 7:00:00 PM 1 record(s) from 7:00:00 PM to 8:00:00 PM 5 record(s) from 8:00:00 PM to 9:00:00 PM 22 record(s) from 9:00:00 PM to 10:00:00 PM 27 record(s) from 10:00:00 PM to 11:00:00 PM 21 record(s) from 11:00:00 PM to 12:00:00 AM
1 6 12 18 24

Observation Type:
2 - DOR
12 - AOR
1 - Basking
15 - Active, off-road
721 - Chorusing

Public Lands Records:
1 - Baker University Wetlands
4 - Clinton Lake
6 - Clinton Lake Project Lands
3 - Clinton Wildlife Area
1 - Copan Wildlife Area
20 - Crawford State Park
4 - Fall River State Park
1 - Fort Leavenworth
1 - Fort Riley
14 - Hillsdale Wildlife Area
1 - Hollister Wildlife Area
15 - Kansas Ecological Reserves
32 - Marais Des Cygnes National Wildlife Refuge
6 - Marais Des Cygnes Wildlife Area
1 - Miami State Fishing Lake and Wildlife Area
6 - Mined Land Wildlife Area
2 - Montgomery State Fishing Lake
5 - Neosho State Fishing Lake
7 - Neosho Wildlife Area
1 - Pomona Lake
3 - Pottawatomie State Fishing Lake No. 1
1 - Sunflower Army Ammunition Plant
6 - Toronto State Park
2 - Tuttle Creek State Parks
6 - Woodson State Fishing Lake and Wildlife Area
GRAY TREEFROG COMPLEX
Hyla chrysoscelis/versicolor, LeConte, 1825/Cope, 1880
  (hI'-la cry-sos'-i-lEs/vers'-i-color)

Recognition:
1-1/4" - 2" (3.2 - 5.1 cm) Rough skin, green to brown, light spot beneath eye, bright yellow or orange on concealed surfaces of hind legs, large toe pads on tips of fingers and toes for climbing vegetation. Few differences exist between this species and the Eastern gray treefrog. Cope's tends to be smaller, has smoother skin. Indistinguishable physically in the field from gray treefrog. Can only be distinguished in the field by their calls.

Confusing Species:
Microscopic inspection of the chromosomes of each species reveals that H. versicolor has an extra set. H. chrysoscelis is diploid, having two complete sets of chromosomes, the usual condition in vertebrates. Hyla versicolor is tetraploid, having double the usual number of chromsomes. Despite currently sharing habitat, the two species generally do not interbreed.

Distribution:
Found throughout the forested regions of the eastern third of Kansas. An isloated record exists at in Pratt County, and may represent an accidental introduction at the state fish hatchery.
Collins and Hillis (1995) and Hillis et al. (1987) analyzed chromosomes of both species throughout Kansas. They found H. chrysoscelis thorughout the eastern third of Kansas, however H. versicolor was found in the Cross Timbers, lower Neoshor River basin, and the Marais des Cygnes and lower Kansas river basins. They determined the specific status for the following specimens:
Hyla chrysoscelis: Chase Co.: KU 203673; Chautauqua Co.: KU 203710; Cherokee Co.: KU 33490, 203665-68, 203706-07, 203720-22; Coffey Co.: KU 203705; Crawford Co.: KU 203718-19; Douglas Co.: KU 16371, 16373-381, 203995-96, 203991-92, 203993-94, 203676- 79, 203712-714; Elk Co.: KU 203711; Franklin Co.: KU 203669-671, 204038; Greenwood Co.: KU 203701; Jackson Co.: KU 203690-94; Jefferson Co.: KU 17398; Johnson Co.: KU 203723-24; Leavenworth Co.: KU 21145; Marshall Co.: KU 49182-84; Miami Co.: KU 203672, 204037; Montgomery Co.: KU 203708-09; Neosho Co.: KU 203715-17; Osage Co.: KU 17681, 203689; Pottawatomie Co. : KU 203674-75; Riley Co. : KU 203698-700; Shawnee Co.: KU 203688; Wabaunsee Co.: KU 203695-97; Wilson Co.: KU 203680-87; Woodson Co.: KU 203702-04
Hyla versicolor: Atchison Co.: KU 203740-41; Bourbon Co.: KU 28672; Cherokee Co.: KU 28671, 31099, 203746; Douglas Co.: KU 8122, 22013, 203745; Elk Co.: KU 203742-44; Franklin Co.: KU 203735; Greenwood Co.: KU 22931; Jefferson Co.: KU 203732-34; Miami Co.: KU 21833, 22112, 203736-39
County Breakdown: County Name (# occurrences)
Allen (15), Anderson (1), Atchison (7), Bourbon (15), Brown (2), Butler (5), Chase (24), Chautauqua (11), Cherokee (203), Clay (9), Coffey (101), Cowley (1), Crawford (29), Dickinson (2), Doniphan (21), Douglas (223), Elk (8), Franklin (30), Geary (2), Greenwood (58), Harper (2), Jackson (56), Jefferson (18), Johnson (8), Labette (35), Leavenworth (66), Linn (78), Lyon (4), Marshall (8), Miami (48), Montgomery (41), Morris (1), Nemaha (10), Neosho (14), Osage (31), Pottawatomie (92), Pratt (1), Republic (1), Riley (10), Shawnee (7), Sumner (5), Wabaunsee (50), Washington (12), Wilson (19), Woodson (20), Wyandotte (3)

Reproduction:
Chorusing is most frequent at night, but individuals often call during daytime in response to thunder or other loud noises. Eggs are laid in batches of ten to forty on the surface of shallow ponds and other small bodies of water. These are usually temporary bodies of water lacking fish. Eggs hatch in approximately five days and metamorphosis takes place at about 45-65 days.

Call Description:
The call of Hyla versicolor is a short high trill, more musical than the Hyla chrysoscelis, and slower. Trill pulse rate for Hyla versicolor is said to vary from 17-35 pulses/second and for Hyla chrysoscelis

Hyla chrysoscelis
<A HREF='calls/395-852.mp3'>[Play Call]</A>
Audio recording by Keith Coleman
Available from Kansas Heritage Photography


Hyla versicolor
<A HREF='calls/Hya_versicolor.mp3'>[Play Call]</A>
Audio recording by Keith Coleman
Available from Kansas Heritage Photography


Growth and Longevity:
KU 21830, Miami County, Ralph Taylor, 8 June 1941, 53 mm (2 1/8 inches), Collins (1993).

Fossil Record:
Pleistocene fossils have been reported from Ellsworth County in central Kansas. This record is outside the currently known range of this species.

Remarks:
Actually a complex of two separate species that are indistinguishable externally. For this reason they are included in the same account, however it should be noted that this arrangement should not minimize their recognition as independent evolutionary entities.

References:
1985. Collins, Joseph T. and David M. Hillis. Final report to the Kansas Fish and Game Commission on the Gray Treefrogs of Kansas. Kansas Fish and Game. Contract #75, 1983-1985: pp. 12.
1987. Hillis, David M., Joseph T. Collins, and James P. Bogart. Distribution of diploid and tetraploid species of Gray Tree Frogs (Hyla chrysoscelis and Hyla versicolor) in Kansas. The American Midland Naturalist. 117(1): pp. 214-217.
2006. Altig, Ronald, Roy W. McDiarmid, Kimberly A. Nichols, and Paul C. Ustach. Tadpoles of the United States and Canada: A Tutorial and Key. Electronic files accessible at http://www.pwrc.usgs.gov/tadpole/. Patuxent Wildlife Research Center, Laurel, MD, USA.. : pp. .
2010. Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. Pp. 400.

User: 54.234.59.94; CCBot/2.0 (http://commoncrawl.org/faq/). © Sternberg Museum of Natural History 1999-2014